In Dark Souls, which boss drops the Lord Soul called the Bequeathed Lord Soul Shard?

In Dark Souls, Gravelord Nito drops the Lord Soul called the Bequeathed Lord Soul Shard.

Nito is one of the four major Lord Soul holders whose defeat is required during the late-game journey to link the First Flame. He is found in the Tomb of the Giants, a dark region reached through the Catacombs. His boss arena contains multiple skeletons, so the battle combines direct combat with the problem of controlling additional enemies.

The Lord Souls are not all obtained in the same way. Seath the Scaleless, the Four Kings, and the Bed of Chaos each provide the other major Lord Soul rewards. Nito’s title, Gravelord, also connects him to the Gravelord Servant covenant, although joining that covenant and defeating Nito are separate parts of the game.