In the 2016 film Assassin's Creed, Callum Lynch uses the Animus to experience Aguilar's memories.
Callum Lynch is a modern-day descendant of the Spanish Assassin Aguilar de Nerha. After being declared dead and rescued from execution, Callum is taken to a facility operated by Abstergo Industries. There, the Animus allows him to access genetic memories inherited through his bloodline.
Michael Fassbender plays Callum Lynch and also portrays Aguilar. These are two distinct characters: Callum lives in the modern period, while Aguilar operates in Spain during the Spanish Inquisition. The film uses their connection to move between the present-day Abstergo plot and Aguilar's historical missions.
Desmond Miles, the protagonist of the early games, does not appear as the film's modern-day lead. Alan Rikkin is an Abstergo executive played by Jeremy Irons, not the character who experiences Aguilar's memories.