Which character is the main villain in Pixar's Cars 2?

Miles Axlerod is the main villain in Pixar's Cars 2.

Axlerod is introduced as a former oil baron who presents the World Grand Prix as a celebration of alternative fuel. Behind that public image, he secretly orchestrates a plot involving Allinol, a fuel that can cause cars to explode when targeted by an electromagnetic weapon. His plan is designed to discredit alternative energy and restore demand for petroleum.

The film follows Lightning McQueen and Mater as they become involved in an international spy adventure. Mater is mistaken for an undercover agent, while British spy Finn McMissile and analyst Holley Shiftwell investigate the conspiracy. Professor Zündapp is an important accomplice, but he is not the mastermind. Francesco Bernoulli is Lightning's Italian racing rival, and Finn McMissile is a hero, making both characters frequent wrong answers to this question.