Which journal published Fleming's 1929 paper on Penicillium cultures?

Fleming’s 1929 paper on Penicillium cultures was published in the British Journal of Experimental Pathology. The article reported the antibacterial action of the mould and introduced the scientific world to his early findings on penicillin.

Fleming reported the work to the journal on 10 May 1929, and the paper appeared in the following month’s issue. Its full title was “On the Antibacterial Action of Cultures of a Penicillium, with Special Reference to their Use in the Isolation of B. influenzae.” Fleming focused heavily on the mould’s usefulness in laboratory culture, especially for isolating bacteria that were not affected by penicillin.

The paper did not immediately trigger the development of a mass-produced medicine. Fleming could not reliably isolate and purify the active substance, and his findings attracted little attention during the 1930s. The Oxford team led by Howard Florey and Ernst Chain revisited the paper in 1939 and transformed the laboratory observation into a practical treatment.

The journal title is sometimes confused with Nature or The Lancet because both published major medical and scientific work related to penicillin later. Fleming’s original 1929 report, however, appeared in the British Journal of Experimental Pathology.